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Williams, B.; Campbell, J.; Kendall, C.; Tuttle, J.; Lynch, E.C., 2023. Exploring the behaviors and social preferences of a large, multigenerational herd of zoo-housed Southern white rhinoceros (Ceratotherium simum simum). Zoo Biology 42 (4): 476-489 - https://doi.org/10.1002/zoo.21758

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Tuttle, J., 2017. Dental surgery for geriatric southern white rhino. Proceedings of the International Rhino Keeper (IRKA) Workshop held in Denver, August 2017 (Abstracts and Powerpoint presentations)

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Tuttle, J., 2015. Which reward (North Carolina Zoo). The Crash April 2015: 2-3

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Tuttle, J., 2014. Rhino open mouth behavior. The Crash January 2014: 6

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Tuttle, J., 2007. Facility focus - the Caldwell Zoo. The Crash 2 (2): 2-3, figs. 1-9

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