 | Holden, M.D.; Gregory, J.; Watkins, V.; Radford, L., 2006. Operant-conditioning programme for white rhinoceros, black rhinoceros and Indian or greater one-horned Asian rhinoceros Ceratotherium simum, Diceros bicornis and Rhinoceros unicornis at Whipsnade Wild Animal Park, Dunstable, UK. International Zoo Yearbook 40: 144-149, pls. 1-2
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