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Gerhardt, U., 1909. Die gegenwartige Stand der Kenntnisse von der Copulationsorgane der Wirbeltiere, insbesondere der Amniota. Ergebnisse und Fortschritte der Zoologie 1: 307-402, figs. 1-16

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Location: World
Subject: Anatomy - Reproductive organs
Species: All Rhino Species


Original text on this topic:
The penis of the Perissodactyla has a great thickness. As in Hyrax and Elephas, the preputial opening in Rhinoceros and Tapirus is located far backwards, and the urine is released backwards. In erection the penis always moves forward. In Rhinoceros (Owen 1868) the penis reaches an enormous size in comparison to the body. In all Perissodactyla a septum in the Corpus fibrosum is absent. The Corpus fibrosum shows a kidney shaped cross-section, as the ventral surface has a space to attach the Corpus spongiosum. The Corpus spongiosum in all genera is well developed and also has a Glans of good proportion, the form of which differs in each genus. In Rhinoceros, the Glans, described by Owen, is extraordinarily large; it forms about a three-sided pyramid, as the basis on the dorsal side has two large and converging excesses. The tip has a very complicated form, the urine conducts lie in a special, well developed Processus urethralis. As general characteristic of the penis of the Perissodactyla we can mention the great size and thickness of the entire organ, the strong development of the Glans and the ansence of a Septum and Penis bones.

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