 | Klaits, B.G., 1973. Upper miocene rhinoceroses from Sansan (Gers), France: the manus. Journal of Paleontology 47 (2): 315-326, pls. 1-5, figs. 1-9
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| Klaits, B.G., 1972. The moving mesaxonic manus - A comparison of tapirs and rhinoceroses. Mammalia 36 (1): 126-145
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