 | Davis, D.D., 1962. Mammals of the lowland rain-forest of North Borneo. Bulletin of the Singapore National Museum 31: 1-129, pls. 1-23, figs. 1-20
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| Davis, D.D., 1958. Mammals of the Kelebit plateau, northern Sarawak. Fieldiana Zoology 39 (15): 119-147, fig. 21, map 1
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