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Richards, D., 1972. Square-lipped rhinoceros: behaviour. Lammergeyer 15: 77-78

Location: Africa - Southern Africa - South Africa
Subject: Behaviour - Fighting
Species: White Rhino

Original text on this topic:
One male and 2 females in the area. On one occasion the male was seen attempting to prevent the females from leaving the area. Each time the females started to move away, the male rounded them up, making a panting sound. The male was also observed spray-urinating. One female appeared to become annoyed at the male and clashed horns.

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